Video - Charles Darwin and the Tree of Life (60mins)
Charles Darwin and the Tree of Life is a 2009 television documentary about Charles Darwin
and his revolutionary theory of evolution through natural selection, produced by the BBC to mark the
bicentenary of Darwin's birth. It is part of the BBC Darwin Season.
The presenter, David Attenborough, outlines the development of the theory by Darwin through his
observations of animals and plants in nature and in the domesticated state, visiting sites important in
Darwin's own life, including Down House, Cambridge University and the Natural History Museum, and using
archive footage from Attenborough's many nature documentaries for the BBC.
He reviews the development of the theory since its beginnings, and its revolutionary impact on the way in
which humans view themselves - not as having dominion over the animals as The Bible says, but as part of
the natural world and subject to the same controlling forces that govern all life on Earth.
Charles Darwin and the Tree of Life - David Attenborough - 60 minutes
OU on the BBC: Charles Darwin And The Tree of Life - About the programme
David Attenborough shares his personal passion for Darwin's work. Find out more about the
programme.
This personal insight into Darwin's 'dangerous idea' from the world's favourite naturalist explores why it is now more important than ever.
Darwin's theory – that life has evolved over millions of years by natural selection – has become so much a 'part of the furniture' in our scientific world that we're in danger of taking it for granted.
But it has never been more important to applaud Darwin's 'unsurpassingly brilliant' insight, or to re-affirm its importance to our entire understanding of how the world works.
David Attenborough is a passionate Darwinian. He sees evolution as the cornerstone of all his thinking; Charles Darwin's ideas have permeated everything David has ever filmed or written.
In this one-hour special, Tree of Life, David asks three key questions: How, and why, did Darwin come up with his theory? Why do we think he was right? And why is it more important now than ever before?
To answer these questions, David takes us on a journey through the last 200 years, tracking the changes in our understanding of the natural world. Even in David's own childhood, there were still many unanswered questions to the theory of evolution.
But since Darwin's time, major scientific discoveries in biology, geology, palaeontology and genetics have all served to underpin and strengthen the evidence for his revolutionary and controversial idea. Today, the pieces of the puzzle fit together so neatly, that there can be little doubt that Darwin was right.
Charles Darwin
From Wikipedia, the free encyclopedia - Text is available under the Creative Commons Attribution-ShareAlike License;
| Charles Darwin | |
|---|---|
Darwin, aged 45 in 1854, by then working towards publication of On the Origin of Species |
|
| Born | Charles Robert Darwin 12 February 1809 The Mount, Shrewsbury, Shropshire, United Kingdom |
| Died | 19 April 1882 (aged 73) Down House, Downe, Kent, United Kingdom |
| Residence | England |
| Citizenship | British |
| Nationality | British |
| Fields | Naturalist |
| Institutions | tertiary education: University of Edinburgh (medicine) University of Cambridge (ordinary Bachelor of Arts) professional institution: Geological Society of London |
| Academic advisors | John Stevens Henslow Adam Sedgwick |
| Known for | The Voyage of the
Beagle On the Origin of Species evolution by natural selection, common descent |
| Influences | Alexander von
Humboldt John Herschel Charles Lyell |
| Influenced | Joseph Dalton Hooker Thomas Henry Huxley George Romanes Ernst Haeckel Sir John Lubbock |
| Notable awards | Royal Medal (1853) Wollaston Medal (1859) Copley Medal (1864) |
| Spouse | Emma Darwin (married 1839) |
Signature
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Darwin published his theory of evolution with compelling evidence in his 1859 book On the Origin of Species, overcoming scientific rejection of earlier concepts of transmutation of species.[3][4] By the 1870s the scientific community and much of the general public had accepted evolution as a fact. However, many favoured competing explanations and it was not until the emergence of the modern evolutionary synthesis from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution.[5][6] In modified form, Darwin's scientific discovery is the unifying theory of the life sciences, explaining the diversity of life.[7][8]
Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh; instead, he helped to investigate marine invertebrates. Studies at the University of Cambridge encouraged his passion for natural science.[9] His five-year voyage on HMS Beagle established him as an eminent geologist whose observations and theories supported Charles Lyell's uniformitarian ideas, and publication of his journal of the voyage made him famous as a popular author.[10]
Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and in 1838 conceived his theory of natural selection.[11] Although he discussed his ideas with several naturalists, he needed time for extensive research and his geological work had priority.[12] He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay which described the same idea, prompting immediate joint publication of both of their theories.[13] Darwin's work established evolutionary descent with modification as the dominant scientific explanation of diversification in nature.[5] In 1871 he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex, followed by The Expression of the Emotions in Man and Animals. His research on plants was published in a series of books, and in his final book, he examined earthworms and their effect on soil.[14]
In recognition of Darwin's pre-eminence as a scientist, he was honoured with a major ceremonial funeral and buried in Westminster Abbey, close to John Herschel and Isaac Newton.[15] Darwin has been described as one of the most influential figures in human history.[16][17]
Life
Childhood and education
See also: Charles Darwin's
education and Darwin-Wedgwood family
Charles Robert Darwin was born in Shrewsbury, Shropshire, England on 12 February
1809 at his family home, The Mount,[18]
He was the fifth of six children of wealthy society doctor and financier Robert Darwin, and Susannah Darwin (née Wedgwood). He
was the grandson of Erasmus Darwin on his father's side, and of
Josiah Wedgwood on his mother's side.
Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire, before going to the University of Edinburgh Medical School with his brother Erasmus in October 1825. He found lectures dull and surgery distressing, so neglected his studies. He learned taxidermy from John Edmonstone, a freed black slave who had accompanied Charles Waterton in the South American rainforest, and often sat with this "very pleasant and intelligent man".[20]
In Darwin's second year he joined the Plinian Society, a student natural history group whose debates strayed into radical materialism. He assisted Robert Edmond Grant's investigations of the anatomy and life cycle of marine invertebrates in the Firth of Forth, and on 27 March 1827 presented at the Plinian his own discovery that black spores found in oyster shells were the eggs of a skate leech. One day, Grant praised Lamarck's evolutionary ideas. Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals. [21] Darwin was rather bored by Robert Jameson's natural history course which covered geology including the debate between Neptunism and Plutonism. He learned classification of plants, and assisted with work on the collections of the University Museum, one of the largest museums in Europe at the time.[22]
This neglect of medical studies annoyed his father, who shrewdly sent him to Christ's College, Cambridge, for a Bachelor of Arts degree as the first step towards becoming an Anglican parson. As Darwin was unqualified for the Tripos, he joined the ordinary degree course in January 1828.[23] He preferred riding and shooting to studying. His cousin William Darwin Fox introduced him to the popular craze for beetle collecting; Darwin pursued this zealously, getting some of his finds published in Stevens' Illustrations of British entomology. He became a close friend and follower of botany professor John Stevens Henslow and met other leading naturalists who saw scientific work as religious natural theology, becoming known to these dons as "the man who walks with Henslow". When his own exams drew near, Darwin focused on his studies and was delighted by the language and logic of William Paley's Evidences of Christianity.[24] In his final examination in January 1831 Darwin did well, coming tenth out of 178 candidates for the ordinary degree.[25]
Darwin had to stay at Cambridge until June. He studied Paley's Natural Theology, which made an argument for divine design in nature, explaining adaptation as God acting through laws of nature.[26] He read John Herschel's new book, which described the highest aim of natural philosophy as understanding such laws through inductive reasoning based on observation, and Alexander von Humboldt's Personal Narrative of scientific travels. Inspired with "a burning zeal" to contribute, Darwin planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. In preparation, he joined Adam Sedgwick's geology course, then travelled with him in the summer for a fortnight, in order to map strata in Wales.[27]
Voyage of the Beagle
For more details on this topic, see Second voyage of HMS
Beagle.
After delays, the voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while the Beagle surveyed and charted coasts.[5][30] He kept careful notes of his observations and theoretical speculations, and at intervals during the voyage his specimens were sent to Cambridge together with letters including a copy of his journal for his family.[31] He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas was a novice and ably collected specimens for expert appraisal.[32] Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with plankton collected in a calm spell.[30][33]
On their first stop ashore at St. Jago, Darwin found that a white band high in the volcanic rock cliffs included seashells. FitzRoy had given him the first volume of Charles Lyell's Principles of Geology which set out uniformitarian concepts of land slowly rising or falling over immense periods,[II] and Darwin saw things Lyell's way, theorising and thinking of writing a book on geology.[34]
When they reached Brazil Darwin was delighted by the tropical forest,[35] but detested the sight of slavery.[36] The survey continued to the south in Patagonia. They stopped at Bahía Blanca, and in cliffs near Punta Alta Darwin made a major find of fossil bones of huge extinct mammals beside modern seashells, indicating recent extinction with no signs of change in climate or catastrophe. He identified the little known Megatherium by a tooth and its association with bony armour which had at first seemed to him like a giant version of the armour on local armadillos. The finds brought great interest when they reached England.[37][38]
On rides with gauchos into the interior to explore geology and collect more fossils, Darwin gained social, political and anthropological insights into both native and colonial people at a time of revolution, and learnt that two types of rhea had separate but overlapping territories.[39][40] Further south he saw stepped plains of shingle and seashells as raised beaches showing a series of elevations. He read Lyell's second volume and accepted its view of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species.[41][42]
Darwin experienced an earthquake in Chile and saw signs that the land had just been raised, including mussel-beds stranded above high tide. High in the Andes he saw seashells, and several fossil trees that had grown on a sand beach. He theorised that as the land rose, oceanic islands sank, and coral reefs round them grew to form atolls.[46][47]
On the geologically new Galápagos Islands Darwin looked for evidence attaching wildlife to an older "centre of creation", and found mockingbirds allied to those in Chile but differing from island to island. He heard that slight variations in the shape of tortoise shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food.[48][49] In Australia the marsupial rat-kangaroo and the platypus seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work.[50] He found the Aborigines "good-humoured & pleasant", and noted their depletion by European settlement.[51]
The Beagle investigated how the atolls of the Cocos (Keeling) Islands had formed, and the survey supported Darwin's theorising.[47] FitzRoy began writing the official Narrative of the Beagle voyages, and after reading Darwin's diary he proposed incorporating it into the account.[52] Darwin's Journal was eventually rewritten as a separate third volume, on natural history.[53]
In Cape Town Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his uniformitarianism as opening bold speculation on "that mystery of mysteries, the replacement of extinct species by others" as "a natural in contradistinction to a miraculous process".[54] When organising his notes as the ship sailed home, Darwin wrote that if his growing suspicions about the mockingbirds, the tortoises and the Falkland Islands Fox were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine".[55] He later wrote that such facts "seemed to me to throw some light on the origin of species".[56]
Inception of Darwin's evolutionary theory
For more details on this topic, see Inception of Darwin's
theory.
Charles Lyell eagerly met Darwin for the first time on 29 October and soon introduced him to the up-and-coming anatomist Richard Owen, who had the facilities of the Royal College of Surgeons to work on the fossil bones collected by Darwin. Owen's surprising results included other gigantic extinct ground sloths as well as the Megatherium, a near complete skeleton of the unknown Scelidotherium and a hippopotamus-sized rodent-like skull named Toxodon resembling a giant capybara. The armour fragments were actually from Glyptodon, a huge armadillo-like creature as Darwin had initially thought.[59][38] These extinct creatures were related to living species in South America.[60]
In mid-December Darwin took lodgings in Cambridge to organise work on his collections and rewrite his Journal.[61] He wrote his first paper, showing that the South American landmass was slowly rising, and with Lyell's enthusiastic backing read it to the Geological Society of London on 4 January 1837. On the same day, he presented his mammal and bird specimens to the Zoological Society. The ornithologist John Gould soon announced that the Galapagos birds that Darwin had thought a mixture of blackbirds, "gros-beaks" and finches, were, in fact, twelve separate species of finches. On 17 February Darwin was elected to the Council of the Geological Society, and Lyell's presidential address presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his uniformitarian ideas.[62]
Early in March, Darwin moved to London to be near this work, joining Lyell's social circle of scientists and experts such as Charles Babbage,[63] who described God as a programmer of laws. Darwin stayed with his freethinking brother Erasmus, part of this Whig circle and close friend of writer Harriet Martineau who promoted Malthusianism underlying the controversial Whig Poor Law reforms to stop welfare from causing overpopulation and more poverty. As a Unitarian she welcomed the radical implications of transmutation of species, promoted by Grant and younger surgeons influenced by Geoffroy. Transmutation was anathema to Anglicans defending social order,[64] but reputable scientists openly discussed the subject and there was wide interest in John Herschel's letter praising Lyell's approach as a way to find a natural cause of the origin of new species.[54]
Gould met Darwin and told him that the Galápagos mockingbirds from different islands were separate species, not just varieties, and what Darwin had thought was a "wren" was also in the finch group. Darwin had not labelled the finches by island, but from the notes of others on the Beagle, including FitzRoy, he allocated species to islands.[65] The two rheas were also distinct species, and on 14 March Darwin announced how their distribution changed going southwards.[66]
Overwork, illness, and marriage
See also: Charles Darwin's health
While developing this intensive study of transmutation, Darwin became mired
in more work. Still rewriting his Journal, he took on editing and publishing the expert
reports on his collections, and with Henslow's help obtained a Treasury grant of £1,000 to sponsor this multi-volume
Zoology of the Voyage of
H.M.S. Beagle, a sum equivalent to about £75,000 in 2011.[68]
He stretched the funding to include his planned books on geology, and agreed unrealistic dates
with the publisher.[69]
As the Victorian era began, Darwin pressed on with
writing his Journal, and in August 1837 began correcting printer's proofs.[70]Darwin's health suffered from the pressure. On 20 September he had "an uncomfortable palpitation of the heart", so his doctors urged him to "knock off all work" and live in the country for a few weeks. After visiting Shrewsbury he joined his Wedgwood relatives at Maer Hall, Staffordshire, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin Emma Wedgwood, nine months older than Darwin, was nursing his invalid aunt. His uncle Jos pointed out an area of ground where cinders had disappeared under loam and suggested that this might have been the work of earthworms, inspiring "a new & important theory" on their role in soil formation which Darwin presented at the Geological Society on 1 November.[71]
William Whewell pushed Darwin to take on the duties of Secretary of the Geological Society. After initially declining the work, he accepted the post in March 1838.[72] Despite the grind of writing and editing the Beagle reports, Darwin made remarkable progress on transmutation, taking every opportunity to question expert naturalists and, unconventionally, people with practical experience such as farmers and pigeon fanciers.[5][73] Over time his research drew on information from his relatives and children, the family butler, neighbours, colonists and former shipmates.[74] He included mankind in his speculations from the outset, and on seeing an orangutan in the zoo on 28 March 1838 noted its childlike behaviour.[75]
On 23 June he took a break and went "geologising" in Scotland. He visited Glen Roy in glorious weather to see the parallel "roads" cut into the hillsides at three heights. He later published his view that these were marine raised beaches, but then had to accept that they were shorelines of a proglacial lake.[77]
Fully recuperated, he returned to Shrewsbury in July. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about career and prospects on two scraps of paper, one with columns headed "Marry" and "Not Marry". Advantages included "constant companion and a friend in old age ... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time."[78] Having decided in favour, he discussed it with his father, then went to visit Emma on 29 July. He did not get around to proposing, but against his father's advice he mentioned his ideas on transmutation.[79]
Malthus and natural selection
Continuing his research in London, Darwin's wide reading now included the sixth edition of Malthus's An Essay on the Principle of Population, and on 28 September 1838 he noted its assertion that human "population, when unchecked, goes on doubling itself every twenty five years, or increases in a geometrical ratio", a geometric progression so that population soon exceeds food supply in what is known as a Malthusian catastrophe. Darwin was well prepared to compare this to de Candolle's "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. He wrote that the "final cause of all this wedging, must be to sort out proper structure, & adapt it to changes", so that "One may say there is a force like a hundred thousand wedges trying force into every kind of adapted structure into the gaps of in the economy of nature, or rather forming gaps by thrusting out weaker ones."[5][80] This would result in the formation of new species.[5][81] As he later wrote in his autobiography;By mid December Darwin saw a similarity between farmers picking the best stock in selective breeding, and a Malthusian Nature selecting from chance variants so that "every part of newly acquired structure is fully practical and perfected",[83] thinking this comparison "a beautiful part of my theory".[84] He later called his theory natural selection, an analogy with what he termed the artificial selection of selective breeding.[5]In October 1838, that is, fifteen months after I had begun my systematic enquiry, I happened to read for amusement Malthus on Population, and being well prepared to appreciate the struggle for existence which everywhere goes on from long-continued observation of the habits of animals and plants, it at once struck me that under these circumstances favourable variations would tend to be preserved, and unfavourable ones to be destroyed. The result of this would be the formation of new species. Here, then, I had at last got a theory by which to work..."[82]
On 11 November, he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters she showed how she valued his openness in sharing their differences, also expressing her strong Unitarian beliefs and concerns that his honest doubts might separate them in the afterlife.[85] While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in Gower Street, then moved his "museum" in over Christmas. On 24 January 1839 Darwin was elected a Fellow of the Royal Society.[86]
On 29 January Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home.[87]
Geology books, Barnacles, evolutionary research
For more details on this topic, see Development of Darwin's
theory.
When FitzRoy's Narrative was published in May 1839, Darwin's Journal and Remarks was such a success as the third volume that later that year it was published on its own.[90] Early in 1842, Darwin wrote about his ideas to Charles Lyell, who noted that his ally "denies seeing a beginning to each crop of species".[91]
Darwin's book The Structure and Distribution of Coral Reefs on his theory of atoll formation was published in May 1842 after more than three years of work, and he then wrote his first "pencil sketch" of his theory of natural selection.[92] To escape the pressures of London, the family moved to rural Down House in September.[93] On 11 January 1844 Darwin mentioned his theorising to the botanist Joseph Dalton Hooker, writing with melodramatic humour "it is like confessing a murder".[94][95] Hooker replied "There may in my opinion have been a series of productions on different spots, & also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject."[96]
Darwin completed his third geological book in 1846. He now renewed a fascination and expertise in marine invertebrates, dating back to his student days with Grant, by dissecting and classifying the barnacles he had collected on the voyage, enjoying observing beautiful structures and thinking about comparisons with allied structures.[101] In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of creation.[102]
In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. James Gully's Malvern spa and was surprised to find some benefit from hydrotherapy.[103] Then in 1851 his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary, and after a long series of crises she died.[104]
In eight years of work on barnacles (Cirripedia), Darwin's theory helped him to find "homologies" showing that slightly changed body parts served different functions to meet new conditions, and in some genera he found minute males parasitic on hermaphrodites, showing an intermediate stage in evolution of distinct sexes.[105] In 1853 it earned him the Royal Society's Royal Medal, and it made his reputation as a biologist.[106] He resumed work on his theory of species in 1854, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in the economy of nature".[107]
Publication of the theory of natural selection
For more details on this topic, see Publication of Darwin's
theory.
Darwin's book was only partly written when, on 18 June 1858, he received a paper from Wallace describing natural selection. Shocked that he had been "forestalled", Darwin sent it on that day to Lyell, as requested by Wallace,[110][111] and although Wallace had not asked for publication, Darwin suggested he would send it to any journal that Wallace chose. His family was in crisis with children in the village dying of scarlet fever, and he put matters in the hands of Lyell and Hooker. After some discussion, they decided on a joint presentation at the Linnean Society on 1 July of On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection; however, Darwin's baby son died of the scarlet fever and he was too distraught to attend.[112]
There was little immediate attention to this announcement of the theory; the president of the Linnean Society remarked in May 1859 that the year had not been marked by any revolutionary discoveries.[113] Only one review rankled enough for Darwin to recall it later; Professor Samuel Haughton of Dublin claimed that "all that was new in them was false, and what was true was old."[114] Darwin struggled for thirteen months to produce an abstract of his "big book", suffering from ill health but getting constant encouragement from his scientific friends. Lyell arranged to have it published by John Murray.[115]
On the Origin of Species proved unexpectedly popular, with the entire stock of 1,250 copies oversubscribed when it went on sale to booksellers on 22 November 1859.[116] In the book, Darwin set out "one long argument" of detailed observations, inferences and consideration of anticipated objections.[117] His only allusion to human evolution was the understatement that "light will be thrown on the origin of man and his history".[118] His theory is simply stated in the introduction:
As many more individuals of each species are born than can possibly survive; and as, consequently, there is a frequently recurring struggle for existence, it follows that any being, if it vary however slightly in any manner profitable to itself, under the complex and sometimes varying conditions of life, will have a better chance of surviving, and thus be naturally selected. From the strong principle of inheritance, any selected variety will tend to propagate its new and modified form.[119]He put a strong case for common descent, and at the end of the book concluded that:
There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved.[120]The last word was the only variant of "evolved" in the first five editions of the book. "Evolutionism" at that time was associated with other concepts, most commonly with embryological development, and Darwin first used the word evolution in The Descent of Man in 1871, before adding it in 1872 to the 6th edition of The Origin of Species.[121]
Responses to publication
For more details on this topic, see Reaction to Darwin's
theory.
The book aroused international interest, with less controversy than had greeted the popular
Vestiges of the
Natural History of Creation.[123]
Though Darwin's illness kept him away from the public debates, he eagerly scrutinised the
scientific response, commenting on press cuttings, reviews, articles, satires and caricatures, and
corresponded on it with
colleagues worldwide.[124]
Darwin had only said "Light will be thrown on the origin of man",[125]
but the first review claimed it made a creed of the "men from monkeys" idea from
Vestiges.[126]
Amongst early favourable responses, Huxley's reviews swiped at Richard Owen, leader of the scientific
establishment Huxley was trying to overthrow.[127]
In April, Owen's review attacked Darwin's friends and condescendingly dismissed his ideas,
angering Darwin,[128]
but Owen and others began to promote ideas of supernaturally guided evolution.[129]The Church of England's response was mixed. Darwin's old Cambridge tutors Sedgwick and Henslow dismissed the ideas, but liberal clergymen interpreted natural selection as an instrument of God's design, with the cleric Charles Kingsley seeing it as "just as noble a conception of Deity".[130] In 1860, the publication of Essays and Reviews by seven liberal Anglican theologians diverted clerical attention from Darwin, with its ideas including higher criticism attacked by church authorities as heresy. In it, Baden Powell argued that miracles broke God's laws, so belief in them was atheistic, and praised "Mr Darwin's masterly volume [supporting] the grand principle of the self-evolving powers of nature".[131] Asa Gray discussed teleology with Darwin, who imported and distributed Gray's pamphlet on theistic evolution, Natural Selection is not inconsistent with Natural Theology.[130][132] The most famous confrontation was at the public 1860 Oxford evolution debate during a meeting of the British Association for the Advancement of Science, where the Bishop of Oxford Samuel Wilberforce, though not opposed to transmutation of species, argued against Darwin's explanation and human descent from apes. Joseph Hooker argued strongly for Darwin, and Thomas Huxley's legendary retort, that he would rather be descended from an ape than a man who misused his gifts, came to symbolise a triumph of science over religion.[130][133]
Even Darwin's close friends Gray, Hooker, Huxley and Lyell still expressed various reservations but gave strong support, as did many others, particularly younger naturalists. Gray and Lyell sought reconciliation with faith, while Huxley portrayed a polarisation between religion and science. He campaigned pugnaciously against the authority of the clergy in education,[130] aiming to overturn the dominance of clergymen and aristocratic amateurs under Owen in favour of a new generation of professional scientists. Owen's claim that brain anatomy proved humans to be a separate biological order from apes was shown to be false by Huxley in a long running dispute parodied by Kingsley as the "Great Hippocampus Question", and discredited Owen.[134]
Darwinism became a movement covering a wide range of evolutionary ideas. In 1863 Lyell's Geological Evidences of the Antiquity of Man popularised prehistory, though his caution on evolution disappointed Darwin. Weeks later Huxley's Evidence as to Man's Place in Nature showed that anatomically, humans are apes, then The Naturalist on the River Amazons by Henry Walter Bates provided empirical evidence of natural selection.[135] Lobbying brought Darwin Britain's highest scientific honour, the Royal Society's Copley Medal, awarded on 3 November 1864.[136] That day, Huxley held the first meeting of what became the influential X Club devoted to "science, pure and free, untrammelled by religious dogmas".[137] By the end of the decade most scientists agreed that evolution occurred, but only a minority supported Darwin's view that the chief mechanism was natural selection.[138]
The Origin of Species was translated into many languages, becoming a staple scientific text attracting thoughtful attention from all walks of life, including the "working men" who flocked to Huxley's lectures.[139] Darwin's theory also resonated with various movements at the time[III] and became a key fixture of popular culture.[IV] Cartoonists parodied animal ancestry in an old tradition of showing humans with animal traits, and in Britain these droll images served to popularise Darwin's theory in an unthreatening way. While ill in 1862 Darwin began growing a beard, and when he reappeared in public in 1866 caricatures of him as an ape helped to identify all forms of evolutionism with Darwinism.[122]
Descent of Man, sexual selection, and botany
- More detailed articles cover Darwin's life from Orchids to Variation, from Descent of Man to Emotions and from Insectivorous Plants to Worms
Enquiries about insect pollination led in 1861 to novel studies of wild orchids, showing adaptation of their flowers to attract specific moths to each species and ensure cross fertilisation. In 1862 Fertilisation of Orchids gave his first detailed demonstration of the power of natural selection to explain complex ecological relationships, making testable predictions. As his health declined, he lay on his sickbed in a room filled with inventive experiments to trace the movements of climbing plants.[140] Admiring visitors included Ernst Haeckel, a zealous proponent of Darwinismus incorporating Lamarckism and Goethe's idealism.[141] Wallace remained supportive, though he increasingly turned to Spiritualism.[142]
The Variation of Animals and Plants under Domestication of 1868 was the first part of Darwin's planned "big book", and included his unsuccessful hypothesis of pangenesis attempting to explain heredity. It sold briskly at first, despite its size, and was translated into many languages. He wrote most of a second part, on natural selection, but it remained unpublished in his lifetime.[143]
His evolution-related experiments and investigations led to books on Insectivorous Plants, The Effects of Cross and Self Fertilisation in the Vegetable Kingdom, different forms of flowers on plants of the same species, and The Power of Movement in Plants. In his last book he returned to The Formation of Vegetable Mould through the Action of Worms.
Death and legacy
In 1882 he was diagnosed with what was called "angina pectoris" which then meant coronary thrombosis and disease of the heart. At the time of his death, the physicians diagnosed "anginal attacks", and "heart-failure".[147]He died at Down House on 19 April 1882. His last words were to his family, telling Emma "I am not the least afraid of death – Remember what a good wife you have been to me – Tell all my children to remember how good they have been to me", then while she rested, he repeatedly told Henrietta and Francis "It's almost worth while to be sick to be nursed by you".[148] He had expected to be buried in St Mary's churchyard at Downe, but at the request of Darwin's colleagues, after public and parliamentary petitioning, William Spottiswoode (President of the Royal Society) arranged for Darwin to be buried in Westminster Abbey, close to John Herschel and Isaac Newton.[15][149]
Children
Darwin's children
|
|
|---|---|
| William Erasmus Darwin | (27 December 1839 – 1914) |
| Anne Elizabeth Darwin | (2 March 1841 – 23 April 1851) |
| Mary Eleanor Darwin | (23 September 1842 – 16 October 1842) |
| Henrietta Emma "Etty" Darwin | (25 September 1843 – 1929) |
| George Howard Darwin | (9 July 1845 – 7 December 1912) |
| Elizabeth "Bessy" Darwin | (8 July 1847 – 1926) |
| Francis Darwin | (16 August 1848 – 19 September 1925) |
| Leonard Darwin | (15 January 1850 – 26 March 1943) |
| Horace Darwin | (13 May 1851 – 29 September 1928) |
| Charles Waring Darwin | (6 December 1856 – 28 June 1858) |
Of his surviving children, George, Francis and Horace became Fellows of the Royal Society,[153] distinguished as astronomer,[154] botanist and civil engineer, respectively. His son Leonard went on to be a soldier, politician, economist, eugenicist and mentor of the statistician and evolutionary biologist Ronald Fisher.[155]
Views and opinions
Religious views
For more details on this topic, see religious views of
Charles Darwin.
By his return he was critical of the Bible as history, and wondered why all religions should not be equally valid.[157] In the next few years, while intensively speculating on geology and transmutation of species, he gave much thought to religion and openly discussed this with Emma, whose beliefs also came from intensive study and questioning.[85] The theodicy of Paley and Thomas Malthus vindicated evils such as starvation as a result of a benevolent creator's laws which had an overall good effect. To Darwin, natural selection produced the good of adaptation but removed the need for design,[158] and he could not see the work of an omnipotent deity in all the pain and suffering such as the ichneumon wasp paralysing caterpillars as live food for its eggs.[132] He still viewed organisms as perfectly adapted, and On the Origin of Species reflects theological views. Though he thought of religion as a tribal survival strategy, Darwin was reluctant to give up the idea of God as an ultimate lawgiver. He was increasingly troubled by the problem of evil.[159][160]
Darwin remained close friends with the vicar of Downe, John Innes, and continued to play a leading part in the parish work of the church,[161] but from around 1849 would go for a walk on Sundays while his family attended church.[156] He considered it "absurd to doubt that a man might be an ardent theist and an evolutionist"[162][163] and, though reticent about his religious views, in 1879 he wrote that "I have never been an atheist in the sense of denying the existence of a God. – I think that generally ... an agnostic would be the most correct description of my state of mind."[85][162]
The "Lady Hope Story", published in 1915, claimed that Darwin had reverted to Christianity on his sickbed. The claims were repudiated by Darwin's children and have been dismissed as false by historians.[164]
Human society
Darwin's views on social and political issues reflected his time and social position. He thought men's eminence over women was the outcome of sexual selection, a view disputed by Antoinette Brown Blackwell in The Sexes Throughout Nature.[165] He valued European civilisation and saw colonisation as spreading its benefits, with the sad but inevitable effect of extermination of savage peoples who did not become civilised. Darwin's theories presented this as natural, and were cited to promote policies which went against his humanitarian principles.[166] Darwin was strongly against slavery, against "ranking the so-called races of man as distinct species", and against ill-treatment of native people.[167][VI]Darwin was intrigued by his half-cousin Francis Galton's argument, introduced in 1865, that statistical analysis of heredity showed that moral and mental human traits could be inherited, and principles of animal breeding could apply to humans. In The Descent of Man Darwin noted that aiding the weak to survive and have families could lose the benefits of natural selection, but cautioned that withholding such aid would endanger the instinct of sympathy, "the noblest part of our nature", and factors such as education could be more important. When Galton suggested that publishing research could encourage intermarriage within a "caste" of "those who are naturally gifted", Darwin foresaw practical difficulties, and thought it "the sole feasible, yet I fear utopian, plan of procedure in improving the human race", preferring to simply publicise the importance of inheritance and leave decisions to individuals.[168] Francis Galton named this field of study "eugenics" in 1883.[V]
Evolutionary social movements
Thomas Malthus had argued that population growth beyond resources was ordained by God to get humans to work productively and show restraint in getting families, this was used in the 1830s to justify workhouses and laissez-faire economics.[169] Evolution was by then seen as having social implications, and Herbert Spencer's 1851 book Social Statics based ideas of human freedom and individual liberties on his Lamarckian evolutionary theory.[170]
Soon after the Origin was published in 1859, critics derided his description of a struggle for existence as a Malthusian justification for the English industrial capitalism of the time. The term Darwinism was used for the evolutionary ideas of others, including Spencer's "survival of the fittest" as free-market progress, and Ernst Haeckel's racist ideas of human development. Writers used natural selection to argue for various, often contradictory, ideologies such as laissez-faire dog-eat dog capitalism, racism, warfare, colonialism and imperialism. However, Darwin's holistic view of nature included "dependence of one being on another"; thus pacifists, socialists, liberal social reformers and anarchists such as Peter Kropotkin stressed the value of co-operation over struggle within a species.[171] Darwin himself insisted that social policy should not simply be guided by concepts of struggle and selection in nature.[172]
After the 1880s a eugenics movement developed on ideas of biological inheritance, and for scientific justification of their ideas appealed to some concepts of Darwinism. In Britain, most shared Darwin's cautious views on voluntary improvement and sought to encourage those with good traits in "positive eugenics". During the "Eclipse of Darwinism" a scientific foundation for eugenics was provided by Mendelian genetics. Negative eugenics to remove the "feebleminded" were popular in America, Canada and Australia, and eugenics in the United States introduced compulsory sterilization laws, followed by several other countries. Subsequently, Nazi eugenics brought the field into disrepute.[V]
The term "Social Darwinism" was used infrequently from around the 1890s, but became popular as a derogatory term in the 1940s when used by Richard Hofstadter to attack the laissez-faire conservatism of those like William Graham Sumner who opposed reform and socialism. Since then it has been used as a term of abuse by those opposed to what they think are the moral consequences of evolution.[173][169]
Commemoration
Main article: Commemoration of Charles
Darwin
More than 120 species and nine genera have been named after Darwin.[177] In one example, the group of tanagers related to those Darwin found in the Galápagos Islands became popularly known as "Darwin's finches" in 1947, fostering inaccurate legends about their significance to his work.[178]
Darwin's work has continued to be celebrated by numerous publications and events. The Linnean Society of London has commemorated Darwin's achievements by the award of the Darwin–Wallace Medal since 1908. Darwin Day has become an annual celebration, and in 2009 worldwide events were arranged for the bicentenary of Darwin's birth and the 150th anniversary of the publication of On the Origin of Species.[179]
Darwin has been commemorated in the UK, with his portrait printed on the reverse of £10 banknotes printed along with a hummingbird and HMS Beagle, issued by the Bank of England.[180]
A life size seated statue of Darwin can be seen in the main hall of the Natural History Museum in London. .[181] A seated statue of Darwin stands in front of Shrewsbury Library, the building that used to house Shrewsbury School, which Darwin attended as a boy.
Darwin College, a postgraduate college at Cambridge University, is named after the Darwin family.
Works
For more details on this topic, see Charles Darwin
bibliography.
Darwin was a prolific writer. Even without publication of his works on evolution, he would have had
a considerable reputation as the author of The Voyage of the Beagle, as a
geologist who had published extensively on South America and had solved the puzzle of the formation
of coral atolls, and as a biologist who had
published the definitive work on barnacles. While On the Origin of Species
dominates perceptions of his work, The Descent of Man and The
Expression of the Emotions in Man and Animals had considerable impact, and his books on
plants including The Power of Movement in
Plants were innovative studies of great importance, as was his final work on The
Formation of Vegetable Mould through the Action of Worms.[182][183]Text is available under the Creative Commons Attribution-ShareAlike License;
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